Article dans une revue: From Thomas Malthus and Pierre Verhulst to Alfred Lotka and Vito Volterra, theoretical biology has studied the dynamics of living species (see Berryman 1992 for an account of this history). The interaction between theoretical biology and game theory (Smith 1982) has also been fruitful, and—as a result of this interaction—a whole discipline of evolutionary game theory has emerged (Weibull 1995).